Sly, N.D., C.R. Freeman-Gallant, A.E., Henschen, P. Minias, L.A. Whittingham, & P.O. Dunn.  2022. Molecular parallelism in signaling function across different sexually selected ornaments in a warbler. Proceedings of the National Academy of Sciences, USA.  119.

He, K., C-H Liang, Y. Zhu, P.O. Dunn, A. Zhao, & P. Minias. 2022. Reconstructing macroevolutionary patterns in avian MHC architecture with genomic data. Frontiers in Genetics.


Minias, P., K. He, & P.O. Dunn 2021. The strength of selection is consistent across both domains of the MHC class I peptide-binding groove in birds. BMC Ecology and Evolution 21: 1-10.

Qin, S., P.O. Dunn, Y. Yang, H. Liu, & K. He. 2021. Polymorphism and varying selection within the MHC class I of four Anas species. Immunogenetics 73: 395-404.

Drzewinska-Chanko, J., R. Wlodarczyk, A. Gajewski, K. Rudnicka, P.O. Dunn, & P. Minias. 2021. Immunocompetent birds choose larger breeding colonies. Journal of Animal Ecology. doi: 10.1111/1365-2656.13540


He, K., P. Minias, & P.O. Dunn 2020. Long-read genome assemblies reveal extraordinary variation in the number and structure of MHC loci in birds. Genome Biology and Evolution, 13(2). doi:10.1093/gbe/evaa270

Minias, P., J. S. Gutierrez & P.O. Dunn. 2020. Avian major histocompatibility complex copy number variation is associated with helminth richness. Biology Letters 16:20200194.

Braun, C.E., P.O. Dunn, G.T. Wann, M.A. Schroeder, & J.W. Hupp. 2020. Body mass and primary molt patterns of greater sage-crouse in Colorado. Western North American Naturalist 80:330-336.


Minias, P., P. O. Dunn, L.A. Whittingham, J.A. Johnson and S. Oyler-McCance. 2019. Evaluation of a Chicken 600K SNP genotyping array in non-model galliform species. Scientific Reports 9: 6407.

Knight, Samantha M., Elizabeth A. Gow, David W. Bradley, Robert G. Clark, Marc Bélisle, Lisha L. Berzins, Tricia Blake, Eli S. Bridge, Lauren Burke, Russell D. Dawson, Peter O. Dunn, Dany Garant, Geoffrey L. Holroyd, David J. T. Hussell, Olga Lansdorp, Andrew J. Laughlin, Marty L. Leonard, Fanie Pelletier, Dave Shutler, Lynn Siefferman, Caz M. Taylor, Helen E. Trefry, Carol M. Vleck, David Vleck, David W. Winkler, Linda A. Whittingham, D. Ryan Norris.  2019. Non-breeding season movements of a migratory songbird are related to declines in resource availability. Auk: Ornithological Advances in press.

Minias P., E. Pikus, L. A. Whittingham and P.O. Dunn. 2019. Evolution of copy number at MHC varies across the avian tree of life. Genome Biology and Evolution 11: 17-28.


Gow, Elizabeth A., L. Burke, D. W. Winkler, S. M. Knight, D. W. Bradley, R. G. Clark, M. Bélisle, L. Berzins, T. Blake, E. S. Bridge, R. D. Dawson, P.O. Dunn, D. Garant, G. Holroyd, A. G. Horn, D. J. T. Hussell, O. Lansdorp, A. J. Laughlin, M. L. Leonard, F. Pelletier, D. Shutler, L. Siefferman, C. M. Taylor, H. Trefry, C. M. Vleck, D. Vleck,  L. A. Whittingham, D. R. Norris. 2018. A range-wide domino effect and resetting of the annual cycle in a migratory songbird. Proceedings of the Royal Society of London B 286: 20181916.

WHITTINGHAM, L.A., P.O. DUNN. C.R. FREEMAN-GALLANT, C.C. TAFF, & J.A. JOHNSON. 2018. Major Histocompatibility Complex variation and blood parasites in resident and migratory populations of the common yellowthroat. Journal of Evolutionary Biology.

KNIGHT, S.M., David W Bradley, Robert G Clark, Elizabeth A Gow, Marc Bélisle, Lisha L Berzins, Tricia Blake, Eli S Bridge, Lauren Burke, Russell D Dawson, Peter O. Dunn, Dany Garant, Geoffrey L Holroyd, David JT Hussell, Olga Lansdorp, Andrew J Laughlin, Marty L Leonard, Fanie Pelletier, Dave Shutler, Lynn Siefferman, Caz M Taylor, Helen E Trefry, Carol M Vleck, David Vleck, David W Winkler, Linda A Whittingham, D Ryan Norris. 2018. Constructing and evaluating a continent-wide migratory songbird network across the annual cycle. Ecological Monographs 88: 445-460.

CLARK, R.D. D.W. WINKLER, R.D. DAWSON, D.J.T. HUSSELL, M.P. LOMBARDO, P.A. THORPE, P.O. DUNN, L.A. WHITTINGHAM. 2018.  Geographic variation and environmental correlates of apparent survival rates in adult tree swallows Tachycineta biolor. Journal of Avian Biology 49: jsv-012514.

AMBARDAR, M., P.O. DUNN, & L.A. WHITTINGHAM. 2018. Reproductive and foraging success of eastern bluebirds (Sialia sialis) in relation to vegetation height. Journal of Field Ornithology 130: 362-370.

MINIAS, P., Z.W. BATESON, L.A. WHITTINGHAM, J.A. JOHNSON, S. OYLER-MCCANCE, & P.O. DUNN. 2018. Extensive shared polymorphism at non-MHC immune genes in recently diverged North American prairie grouse. Immunogenetics 70: 195-204. DOI 10.1007/s00251-017-1024-4

MINIAS, P., E. PIKUS, L.A. WHITTINGHAM, & P.O. DUNN. 2018. A global analysis of selection at the avian MHC. Evolution 72: 1278-1293.

HENSCHEN, A.E., L.A. WHITTINGHAM, & P.O. DUNN. 2018. Male stress response is related to ornamentation but not resistance to oxidative stress in a warbler. Functional Ecology 32: 1810-1818.


HENSCHEN, A.E., L.A. WHITTINGHAM, & P.O. DUNN. 2017. The relationship between blood parasites and ornamentation depends on the level of analysis in the common yellowthroat. Journal of Avian Biology 48: 1263-1272.

MINIAS, P., L.A. WHITTINGHAM, & P.O. DUNN. 2017. Coloniality and migration are related to selection on MHC genes in birds. Evolution 71: 432-441. DOI: 10.1111/evo.13142


WHITTINGHAM, L. A. & P.O. DUNN. 2016. Experimental evidence that brighter males sire more extra-pair young in tree swallows. Molecular Ecology 25: 3706-3715.

BATESON, Z.W., S.C. HAMMERLY, J.A. JOHNSON, M.E. MORROW, L.A. WHITTINGHAM, & P.O. DUNN. 2016. Specific alleles at immune genes, rather than genome-wide heterozygosity, are related to immunity and survival in the critically endangered Attwater’s prairie-chicken. Molecular Ecology 25: 4730–4744.

MINIAS, P., Z.W. BATESON, L.A. WHITTINGHAM, J.A. JOHNSON, S. OYLER-MCCANCE, & P.O. DUNN. 2016. Contrasting evolutionary histories of MHC class I and class II loci in grouse – effects of selection and gene conversion. Heredity 116: 466-476. DOI: 10.1038/hdy.2016.6

HENSCHEN, A.E., L.A. WHITTINGHAM, & P.O. DUNN. 2016. Oxidative stress is related to both melanin and carotenoid-based ornaments in the common yellowthroat. Functional Ecology 30: 749-758.  DOI: 10.1111/1365-2435.12549.


BATESON, Z.W., L.A. WHITTINGHAM, J.A. JOHNSON, & P.O. DUNN. 2015. Contrasting patterns of selection and drift between two categories of immune genes in prairie-chickens. Molecular Ecology 24: 6095-6106.  

FAIRHURST, G. D., L. L. BERZINS, D. W. BRADLEY, A. J. LAUGHLIN, A. ROMANO, M. ROMANO, C. SCANDOLARA, R. AMBROSINI, R. D. DAWSON, P.O. DUNN, K. A. HOBSON, F. LIECHTI, T. A. MARCHANT, D. R. NORRIS, D. RUBOLINI, N. SAINO, C. M. TAYLOR, L. A. WHITTINGHAM, & R. G. CLARK. 2015. Assessing costs of carrying geolocators using feather corticosterone in two species of aerial insectivore. Royal Society Open Science 2: 150004

DUNN, P.O., J.K. ARMENTA, & L.A. WHITTINGHAM. 2015. Natural and sexual selection act on different axes of variation in avian plumage color. Science Advances 1 (2): 1400155.

WHITTINGHAM, L.A., C.R. FREEMAN-GALLANT, C.C. TAFF, & P.O. DUNN. 2015. Different ornaments signal male health and MHC variation in two populations of a warbler. Molecular Ecology 24: 1584-1595.


BATESON, Z.W., P.O. DUNN, S.D. HULL, A.E. HENSCHEN, J.A. JOHNSON, & L.A. WHITTINGHAM. 2014. Genetic restoration of a threatened population of greater prairie-chickens. Biological Conservation 174: 12-19.

WHITTINGHAM, L. A. & P.O. DUNN.  2014. Extra-pair mating and sexual selection on male traits across populations. Wilson Journal of Ornithology 126: 9-18.

WINKLER, D.W., K.M. RINGELMAN, P.O. DUNN, L.A. WHITTINGHAM, D. J.T. HUSSELL, R.G. CLARK, R.D. DAWSON, L.S. JOHNSON, A. ROSE, S. H. AUSTIN, W.D. ROBINSON, M.P. LOMBARDO, P.A. THORPE, D. SHUTLER, R.J. ROBERTSON, M. STAGER, M. LEONARD, A. G. HORN, J. DICKINSON, V. FERRETTI, V. MASSONI, F. BULIT, J.C. REBOREDA, M. LILJESTHRÖM, M. QUIROGA, E. RAKHIMBERDIEV, & D.R. ARDIA. 2014. Latitudinal variation in clutch size–lay date regressions in Tachycineta swallows: effects of food supply or demography? Ecography 37:670-678.

BRADLEY, D.W., R.G. CLARK, P.O. DUNN, A. J. LAUGHLIN, C.M. TAYLOR, C. VLECK, L.A. WHITTINGHAM, D.W. WINKLER, & D.R. NORRIS. 2014. Trans-Gulf of Mexico loop migration of tree swallows revealed by solar geolocation. Current Zoology 60: 653-659.

FREEMAN-GALLANT, C.R., R.L. SCHNEIDER, C.C. TAFF, P.O. DUNN, & L.A. WHITTINGHAM. 2014. Contrasting patterns of selection on the size and coloration of a female plumage ornament in common yellowthroats. Journal of Evolutionary Biology 27: 982-991.

DUNN, P.O., & A.P. MOLLER. 2014. Changes in breeding phenology and population size of birds. Journal of Animal Ecology 83:729-739


SEDDON, N., C.A. BOTERO, J.A. TOBIAS, P.O. DUNN, H.E.A. MACGREGOR, D.R. RUBENSTEIN, J.A.C. UY, J.T. WEIR, L.A. WHITTINGHAM, & R.J. SAFRAN. 2013. Sexual selection accelerates signal evolution during speciation in birds. Proceedings of the Royal Society of London B 280: 1471-2954.

LAUGHLIN, A. J., C. M. TAYLOR, D.W. BRADLEY, D. LECLAIR, R.C. CLARK, R.D. DAWSON, P.O. DUNN, A. HORN, M. LEONARD, D. R. SHELDON, D. SHUTLER, L. A. WHITTINGHAM, D. W. WINKLER, & D. R. NORRIS. 2013. Integrating information from geolocators, weather radar, and citizen science to uncover a key stopover area of an aerial insectivore. The Auk 130: 230-239.

DUNN, P.O., J.L. BOLLMER, C.R. FREEMAN-GALLANT & L.A. WHITTINGHAM. 2013. MHC variation is related to a sexually selected ornament, survival, and parasite resistance in the common yellowthroat. Evolution 67: 679-687. (PDF)

TAFF, C.C. , C.R. FREEMAN-GALLANT, P.O. DUNN & L.A. WHITTINGHAM. 2013. Spatial distribution of nests constrains the strength of sexual selection in a warbler. Journal of Evolutionary Biology 26: 1392-1405.

EIMES, J.A., K. REED, K. MENDOZA, J.L. BOLLMER, L.A. WHITTINGHAM, Z.W. BATESON & P.O. DUNN. 2013. Greater prairie-chickens have a compact MHC-B with a single class IA locus. Immunogenetics 65: 133-144. (PDF)


BOLLMER, J.L., P.O. DUNN, C.R. FREEMAN-GALLANT & L.A. WHITTINGHAM. 2012. Social and extra-pair mating in relation to MHC variation in common yellowthroats. Proceedings of the Royal Society of London B 279: 4778-4785. (PDF)

TAFF, C.C. , C.R. FREEMAN-GALLANT, P.O. DUNN & L.A. WHITTINGHAM. K. BELINSKY, D STEINBERGER, C. CLARK, & H. SACKS. 2012. Multi-modal sexual selection in a warbler: plumage and song are related to different fitness components. Animal Behaviour 84: 813-821.

SHUTLER, D. D. HUSSELL, R. NORRIS, D. WINKLER, R. ROBERTSON, F. BONIER, W. RENDELL, M. BÉLISLE, R. CLARK, R. DAWSON, N. WHEELWRIGHT, M. LOMBARDO, P. THORPE, M. TRUAN, R. WALSH, M. LEONARD, A. HORN, C. VLECK, D. VLECK, A. ROSE, L. WHITTINGHAM, P. DUNN, K. HOBSON, & M. STANBACK. 2012. Spatiotemporal Patterns in Nest Box Occupancy by Tree Swallows Across North America. Avian Conservation and Ecology 7 (1): 3. (PDF)

HESS, B.D., P.O. DUNN & L.A. WHITTINGHAM. 2012. Females choose multiple mates in the lekking greater prairie-chicken. Auk 129: 133-139. (PDF)


EIMES, J.A., J.L. BOLLMER, L.A. WHITTINGHAM, J.A. JOHNSON, C. VAN OOSTERHOUT & P.O. DUNN. 2011. Rapid loss of MHC class II variation in a bottlenecked population is explained by drift and loss of copy number variation. Journal of Evolutionary Biology 24: 1847-1856. (PDF)

BOLLMER, J.L., E.A. RUDER, J.A. JOHNSON, J.A. EIMES & P.O. DUNN. 2011. Drift and selection influence geographic variation at immune loci of prairie-chickens. Molecular Ecology 20: 4695–4706. (PDF)

TAFF, C.C., C.R. FREEMAN-GALLANT, P.O. DUNN & L.A. WHITTINGHAM. 2011. Relationship between brood sex ratio and male ornaments depends on male age in a warbler. Animal Behaviour 81: 619-625. (PDF)

DUNN, P.O., D.W. WINKLER, L.A. WHITTINGHAM, S.J. HANNON, & R.J. ROBERTSON. 2011. A test of the mismatch hypothesis: how predictable is the timing of food abundance for aerial insectivores? Ecology 92: 450-461. (PDF)


DUNN, P.O., & D.W. WINKLER. 2010. Effects of climate change on timing of breeding and reproductive success in birds. Chapter 10 In: Effects of climate change on birds. A. P. Møller, W. Fiedler and P. Berthold (eds.). Oxford Univ. Press. (email for PDF)

WHITTINGHAM, L.A., & P.O. DUNN. 2010. Fitness benefits of polyandry for experienced females. Molecular Ecology 19:2328-2335. (PDF)

BOLLMER, J.L. , P.O. DUNN, L.A. WHITTINGHAM, & C. WIMPEE. 2010. Extensive MHC class II B gene duplication in a passerine, the Common Yellowthroat (Geothlypis trichas). Journal of Heredity 101: 448-460. (PDF)

FREEMAN-GALLANT, C.R., C.C. TAFF, D.F. MORIN, P.O. DUNN, L.A. WHITTINGHAM, & S.M. TANG. 2010. Sexual selection, multiple ornaments, and age- and condition-dependent signalling in the common yellowthroat. Evolution. 64: 1007-1017. (PDF)

EIMES, J.A., J.L. BOLLMER, P.O. DUNN. L.A. WHITTINGHAM, & C. WIMPEE. 2010. MHC class II diversity and balancing selection in greater prairie-chickens. Genetica 138: 265-271. (PDF)

DUNN. P.O., J. GARVIN, L.A. WHITTINGHAM, C. FREEMAN-GALLANT, & D. HASSELQUIST. 2010. Carotenoid and melanin-based ornaments signal similar aspects of male quality in two populations of the common yellowthroat. Functional Ecology 24:149-158. (PDF)


DUNN. P.O., J.T. LIFJELD, & L.A. WHITTINGHAM. 2009. Multiple paternity and nestling quality in tree swallows. Behavioral Ecology and Sociobiology 63:911-922. (PDF)


ARMENTA, J., P.O. DUNN, & L.A. WHITTINGHAM. 2008. Effects of specimen age on plumage color. Auk 125: 803-808. (PDF)

ARMENTA, J., P.O. DUNN, & L.A. WHITTINGHAM. 2008. Quantifying avian sexual dichromatism: a comparison of methods.
Journal of Experimental Biology 211:2423-2430 (PDF)

DUNN, P.O., L.A. WHITTINGHAM, C. FREEMAN-GALLANT, & J. DeCOSTE. 2008. Geographic variation in the function of ornaments in the common yellowthroat. Journal of Avian Biology 39:66-72. (PDF)

GARVIN, J., P.O. DUNN, L.A. WHITTINGHAM, D. STEEBER, & D. HASSELQUIST. 2008. Do male ornaments signal immunity in common yellowthroats? Behavioral Ecology 19: 54-60 (PDF)


DALE, J., P.O. DUNN, J. FIGUEROLA, T. LISLEVAND, T. SZEKELY, & L.A. WHITTINGHAM. 2007. Sexual selection explains Rensch’s rule of allometry for sexual size dimorphism. Proceedings of the Royal Society of London B 274: 2971-2979. (PDF)

WHITTINGHAM, L.A., P.O. DUNN, & J.T. LIFJELD. 2007. Egg size is related to nestling quality, but there is no differential allocation in tree swallows. Condor 109: 585-594. (PDF)

JOHNSON, J., P.O. DUNN, & J. BOUZAT. 2007. Effects of recent population bottlenecks on reconstructing the demographic history of prairie-chickens. Molecular Ecology 16: 2203-2222. (PDF)

MITCHELL, D.P., P.O. DUNN, L.A. WHITTINGHAM, & C. FREEMAN-GALLANT. 2007. Attractive males provide less parental care in two populations of the common yellowthroat. Animal Behaviour 73: 165-170 (PDF)

ABROE, B., J. GARVIN, M. PEDERSEN, L.A. WHITTINGHAM, & P.O. DUNN. 2007. Brood sex ratios are related to male size, but not attractiveness in the common yellowthroat. Auk 124: 176-184. (PDF).

DUNN. P.O., & L.A. WHITTINGHAM. 2007. Search costs influence the spatial distribution, but not the level of extra-pair mating in tree swallows. Behavioral Ecology and Sociobiology 61: 449-454. (PDF)


PEDERSEN, M., P.O. DUNN, & L.A. WHITTINGHAM. 2006. Extra-territorial forays are related to a male ornamental trait in common yellowthroats. Animal Behaviour 72: 479-486. (PDF)

GARVIN, J., B. ABROE, B., M. PEDERSEN, P.O. DUNN & L.A. WHITTINGHAM. 2006. Immune response of nestling warblers varies with extra-pair paternity and temperature. Molecular Ecology 15: 3833-3840. (PDF)

JOHNSON, J., & P.O. DUNN. 2006. Low genetic variation in the heath hen prior to extinction and implications for the conservation of prairie-chicken populations. Conservation Genetics 7: 37-48. (PDF)

WHITTINGHAM, L.A., P.O. DUNN, & STAPLETON, M. 2006. Repeatability of extra-pair mate choice by female tree swallows. Molecular Ecology 15: 841-849. (PDF)


DUNN, P.O. & L.A. WHITTINGHAM. 2005. Radio-tracking of female tree swallows prior to egg-laying. Journal of Field Ornithology 76: 259-263. (PDF)

NOOKER, J., P.O. DUNN, & L.A. WHITTINGHAM. 2005. Effects of food abundance, weather and female condition on reproduction in tree swallows. Auk 122: 1225-1238. (PDF)

TORTI, V. & P.O. DUNN. 2005. Variable effects of climate change on six species of North American birds. Oecologia 145: 486-495. (PDF)

WHITTINGHAM, L.A., P.O. DUNN, & NOOKER, J. 2005. Maternal influences on brood sex ratios: an experimental study in tree swallows. Proceedings of the Royal Society of London B 272: 1775-1780. (PDF)

TAROF, S., P.O. DUNN & L. A. WHITTINGHAM. 2005. Dual functions of a melanin-based ornament in the common yellowthroat. Proceedings of the Royal Society of London B 272: 1121-1127. (PDF)

PITCHER, T., P.O. DUNN, & L.A. WHITTINGHAM. 2005. Sperm competition and the evolution of avian testes size. Journal of Evolutionary Biology 18: 557-567. (PDF)

WHITTINGHAM, L.A. & P.O. DUNN. 2005. Effects of extra-pair and within-pair reproductive success on the opportunity for selection in birds. Behavioral Ecology 16: 138-144. (PDF)


DUNN, P.O. 2004. Breeding dates and reproductive performance. Chapter 4 In: Birds and climate change. A. P. Møller, W. Fiedler and P. Berthold (eds.). Academic Press, London. Advances in Ecological Research 35: 69-87.

JOHNSON, J., R. BELLINGER, J. TOEPFER & P.O. DUNN. 2004. Temporal changes in allele frequencies and low effective population size in greater prairie-chickens. Molecular Ecology 13: 2617-2630 (PDF).

POIRIER, N.E., L.A. WHITTINGHAM & P.O. DUNN. 2004. Males achieve greater reproductive success through multiple broods than through extra-pair mating in house wrens. Animal Behaviour 67: 1109-1116 (PDF).

SPARKS, T.H., H.Q.P. CRICK, P.O. DUNN, & L.V. SOKOLOV. 2004. Phenology of selected lifeforms: Birds. In: Phenology: An integrative environmental science, M.D. Schwartz (Ed.), pp 421-436. Kluwer, Amsterdam.


JOHNSON, J., J. TOEPFER & P.O. DUNN. 2003. Contrasting patterns of mitochondrial and microsatellite population structure in fragmented populations of greater prairie chickens. Molecular Ecology 12: 3335-3347 (PDF).

POIRIER, N.E., L.A. WHITTINGHAM & P.O. DUNN. 2003. Effects of paternity and mate availability on mate switching in house wrens. Condor 105: 816-821 (PDF)

BELLINGER, R., J. JOHNSON, J. TOEPFER, & P.O. DUNN. 2003. Loss of genetic variation in greater prairie chickens following a population bottleneck in Wisconsin, USA. Conservation Biology 17: 717-724 (PDF).

WHITTINGHAM, L.A., P.O. DUNN & E. CLOTFELTER. 2003. Parental allocation of food to nestling tree swallows: the influence of nestling behavior, sex, and paternity. Animal Behaviour 65: 1203-1210 (PDF).

GALEOTTI, P., D. RUBOLINI, P.O. DUNN, & M. FASOLA. 2003. Colour polymorphism in birds: causes and functions. Journal of Evolutionary Biology 16: 635-646. (PDF)


WINKLER, D.W., P.O. DUNN, & C.E. McCULLOCH. 2002. Predicting the effects of climate change on avian life-history traits. Proceedings of the National Academy of Sciences, USA 99: 13595-13599 (PDF).

LIFJELD, J., P.O. DUNN, & L.A. WHITTINGHAM. 2002. Energetic stress, immunosuppression and parental effort in tree swallows. Oecologia 130: 185-190 (PDF).

WHITTINGHAM, L.A., S.M. VALKENAAR, N.E. POIRIER & P.O. DUNN. 2002. Maternal condition and nestling sex ratio in House Wrens. Auk 119: 125-13 (PDF).


BADYAEV, A.V., G.E. HILL, P.O. DUNN, & J.C. GLEN. 2001. Plumage color as a composite trait: functional integration of sexual ornamentation in the house finch. American Naturalist 158: 221-235 (PDF).

STAPLETON, M., P.O. DUNN, J. McCARTY, A. SECORD & L.A. WHITTINGHAM. 2001. Polychlorinated Biphenyl contamination and minisatellite DNA mutation rates of tree swallows. Environmental Toxicology and Chemistry 20: 2263-2267 (PDF).

THUSIUS, K., K. PETERSON, P.O. DUNN, & L.A. WHITTINGHAM. 2001. Male mask size is correlated with mating success in common yellowthroats. Animal Behaviour 62: 435-446 (PDF).

THUSIUS, K., P.O. DUNN, K. PETERSON, & L.A. WHITTINGHAM. 2001. Extrapair paternity is influenced by breeding synchrony and density in the common yellowthroat. Behavioral Ecology 12:633-639 (PDF).

WHITTINGHAM, L.A. & P. O. DUNN. 2001. Survival of withinpair and extrapair young in tree swallows. Behavioral Ecology 12: 496-500 (PDF)

PETERSON, K., K. THUSIUS, L.A. WHITTINGHAM, & P.O. DUNN. 2001. Allocation of male parental care in relation to paternity within and among broods of common yellowthroats. Ethology 107: 573-586 (PDF).

DUNN. P.O., L. A. WHITTINGHAM, & T. PITCHER. 2001. Mating systems, sperm competition and the evolution of sexual dimorphism in birds. Evolution 55: 161-175 (PDF).

WHITTINGHAM, L.A. & P.O. DUNN. 2001. Paternity and male parental care in birds. Current Ornithology 16: 257-298. (PDF)


WHITTINGHAM, L.A. & P.O. DUNN. 2000. Offspring sex ratios in tree swallows: females in better condition produce more sons. Molecular Ecology 9: 1123-1129 (PDF).

CLOTFELTER, E., WHITTINGHAM, L.A. & P. O. DUNN. 2000. Laying order, hatching asynchrony, and nestling survival in tree swallows. Journal of Avian Biology 31: 329-334. (PDF)

HEINSOHN, R., P. DUNN, S. LEGGE & M. DOUBLE. 2000. Coalitions of relatives and reproductive skew in cooperatively breeding white-winged choughs. Proceedings of the Royal Society of London B 267: 243-249. (PDF)

DUNN, P.O., K. THUSIUS, D. WINKLER, & K. KIMBER. 2000. Geographic and ecological variation in clutch size of tree swallows. Auk 117: 215-221. (PDF)


DUNN, P.O., & D.W. WINKLER. 1999. Climate change has affected the breeding date of tree swallows throughout North America. Proceedings of the Royal Society of London B. 266: 2487-2490. (PDF)

HEINSOHN, R., S. LEGGE, & P.O. DUNN. 1999. Extreme reproductive skew in cooperatively-breeding birds: tests of theory in white-winged choughs. Pp. 2858-2878 in Adams, N.J. & Slotow, R.H. eds. Proceedings of the 22nd International Ornithological Congress, BirdLife South Africa, Johannesburg.

DUNN, P.O., & A. COCKBURN. 1999. Extra-pair mate choice and honest signaling in cooperatively-breeding superb fairy-wrens. Evolution 53: 938-946. (PDF)

DUNN, P.O., A. AFTON, M. GLOUTNEY, & R.T. ALISAUSKAS. 1999. Forced copulation results in few extra-pair fertilizations in Ross’s and lesser snow geese. Animal Behaviour 57: 1071-1081.


WHITTINGHAM, L.A., & P.O. DUNN. 1998. Male parental effort and paternity in a variable mating system. Animal Behaviour 55: 629-640. (PDF)

DUNN, P.O., & A. COCKBURN. 1998. The costs and benefits of extra-group mating in superb fairy-wrens. Pages 147-162 in P. Parker, N. Burley (eds.), Ornithological Monograph No. 49, Avian Reproductive Tactics: Female and Male Perspectives. Amer. Ornithol. Union, Washington, D.C.


WHITTINGHAM, L.A., P.O. DUNN, & R. MAGRATH. 1997. Relatedness, polyandry and extra-group paternity in the cooperatively breeding white-browed scrubwren (Sericornis frontalis). Behavioral Ecology and Sociobiology 40: 261-270.


DUNN, P.O. & A. COCKBURN. 1996. Evolution of male parental care in a bird with almost complete cuckoldry. Evolution 50: 2542-2548. (PDF)


GREEN, D., A. COCKBURN, M. HALL, H. OSMOND, & P.O. DUNN. 1995. The tradeoff between mating and parental effort in male superb fairy-wrens. Proceedings of the Royal Society, London B. 262:297-303.

P.O. DUNN, P. O., A. COCKBURN, & R. A. MULDER. 1995. Fairy-wren helpers often care for young to which they are unrelated. Proceedings of the Royal Society, London B. 259: 339-343.

WHITTINGHAM, L. A., P.O. DUNN, & R. J. ROBERTSON. 1995. Testing the female mate guarding hypothesis. Animal Behaviour 50: 277-279.


DUNN, P. O., R. J. ROBERTSON, D. MICHAUD-FREEMAN & P. T. BOAG. 1994. Extra-pair paternity in tree swallows: why do females mate with more than one male? Behavioral Ecology and Sociobiology 35: 273-281.

WHITTINGHAM, L. A., P.O. DUNN, & R. J. ROBERTSON. 1994. Female response to reduced male parental care in birds: an experiment in tree swallows. Ethology 96: 260-269.

LIFJELD, J. T., P.O. DUNN, & D. F. WESTNEAT. 1994. Sexual selection by sperm competition in birds: male-male competition or female choice? Journal of Avian Biology 25: 244-250.

DUNN, P.O., L. A. WHITTINGHAM, J. T. LIFJELD, R. J. ROBERTSON, & P. T. BOAG. 1994. Effects of breeding density, synchrony and experience on extra-pair paternity in tree swallows. BehavioralEcology 5: 123-129.

MULDER, R. A., P.O. DUNN, A. COCKBURN, K. LAZENBY-COHEN, & M. J.HOWELL. 1994. Helpers liberate female fairy-wrens from constraints on extra-pair mate choice. Proceedings of the Royal Society, London B 255: 223-229.

DUNN, P.O. & J.T. LIFJELD. 1994. Can extra-pair copulations be used to predict extra-pair paternity? Animal Behaviour 47: 983-985.

WHITTINGHAM, L. A., P.O. DUNN, & R. J. ROBERTSON. 1994. Do female tree swallows guard their mates by copulating frequently? Animal Behaviour 47: 994-997.


WHITTINGHAM, L.A., P.O. DUNN, & R.J. ROBERTSON. 1993. Confidence of paternity and male parental care: an experimental study in tree swallows. Animal Behaviour 46: 139-147.

LIFJELD, J.T., P.O. DUNN, R.J. ROBERTSON, & P.T. BOAG. 1993. Extra-pair paternity in monogamous tree swallows. Animal Behaviour 45: 213-229.

DUNN, P.O., & R.J. ROBERTSON. 1993. Extra-pair paternity in polygynous tree swallows. Animal Behaviour 45: 231-239.

VENIER, L.A., P.O. DUNN, J.T. LIFJELD, & R.J. ROBERTSON. 1993. Behavioural patterns of extra-pair copulation in tree swallows. Animal Behaviour 45: 412-415.


DUNN, P.O., & R.J. ROBERTSON. 1992. Geographic variation in the importance of male parental care and mating systems in tree swallows. BehavioralEcology 3: 291-299.

DUNN, P.O., & S.J. HANNON. 1992. The effects of food abundance and male parental care on reproductive success and monogamy in tree swallows. Auk 109: 488-499.

DUNN, P. 1992. Do male birds adjust territory size to the risk of cuckoldry? Animal Behaviour 43: 857-859.


DUNN, P.O., & S.J. HANNON. 1991. Intraspecific competition and the maintenance of monogamy in tree swallows. Behavioral Ecology 2: 258-266.


DUNN, P.O., & S.J. HANNON. 1989. Evidence for obligate male parental care in black-billed magpies. Auk 106: 635-644.

DUNN, P.O., T.A. MAY, M.A. MCCOLLOUGH, & M.A. HOWE. 1988. Length of stay and fat content of migrant semipalmated sandpipers in eastern Maine. Condor 90: 824-835.

DUNN, P.O., & C.E. BRAUN. 1986. Summer habitat use by adult female and juvenile sage grouse. Journal of Wildlife Management 50: 228-235.

DUNN, P.O. & R.A. RYDER. 1986. Notes on the birds of Cold Spring Mountain, northwestern Colorado. Great BasinNaturalist 46: 651-655.

DUNN, P.O., & C.E. BRAUN. 1986. Late summer-spring movements of juvenile sage grouse. Wilson Bulletin 98: 83-92.

DUNN, P.O., & C.E. BRAUN. 1985. Natal dispersal and lek fidelity of sage grouse. Auk 102: 621-627.